There’s an old saw in philanthropy: If you’ve seen one foundation, you’ve seen one foundation. Each is distinctive, which makes sense: Extremely wealthy people do not get to be that way by following the crowd, so they want their foundations to stand out as well.

Still, of the 86,000 or so grantmaking foundations in the United States, few stand quite so far outside of the mainstream as the Open Philanthropy Project, which guides the charitable giving of Dustin Moskovitz, the cofounder of...


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